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Rusty (Hatosy) starts to pursue a path to a more meaningful life, thanks to his connection to Bob (Goldblum), the boyfriend of his mother, Mary (Lynch). His new take on life causes friction with his best friend, Dallas (Cann), and both men find their friendship pushed to its breaking point, causing them to make life-changing decisions.
Internationally famous oceanographer Steve Zissou (Bill Murray) and his crew—Team Zissou—set sail on an expedition to hunt down the mysterious, elusive, possibly non-existant Jaguar Shark that killed Zissou's partner during the documentary filming of their latest adventure. They are joined on their voyage by a young airline co-pilot who may or may not be Zissou's son (Owen Wilson), a beautiful journalist (Cate Blanchett) assigned to write a profile of Zissou, and Zissou's estranged wife and co-producer, Eleanor (Anjelica Huston). They face overwhelming complications including pirates, kidnapping, and bankruptcy.