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The sleepy town of Wheelsy could be any small town in America – somewhat quaint and gentle, peopled with friendly folks who mind their own business. But just beneath the surface charm, something unnamed and evil has arrived…and is growing. No one seems to notice as telephone poles become clogged with missing pet flyers, or when one of the town's richest citizens, Grant Grant (Michael Rooker), begins to act strangely. But when farmers' livestock turn up horribly mutilated and a young women goes missing, Sheriff Bill Pardy (Nathan Fillion) and his team, aided by Grant's wife Starla (Elizabeth Banks), uncover the dark force laying siege to their town… and come face-to-face with an older-than-time organism intent on absorbing and devouring all life on Earth.
Devastated when his fiancée Kate (Eva Longoria Parker) is killed on their wedding day, Henry (Paul Rudd) reluctantly agrees to consult a psychic named Ashley (Lake Bell) at the urging of his sister Chloe (Lindsay Sloane). Despite his skepticism over her psychic abilities, Henry finds himself falling hard for Ashley, and vice versa. But there is a big snag. Ashley is being haunted by Kate's ghost, who considers it her heavenly duty to break up Henry and Ashley's fledgling romance, if it is the last thing she does on this earthly plane...