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Ben Stiller portrays risk-averse Reuben Feffer, whose best-laid plans for life and love careen wildly off track when his bride (Debra Messing) dumps him on their honeymoon for a muscle-bound scuba instructor (Hank Azaria). Stunned, humiliated and in the grip of acute indigestion, Reuben plans to play it safer than ever. But a chance encounter with an adventure-craving, childhood friend named Polly (Jennifer Aniston) shoots him into a whirlwind of extreme sports, spicy foods, ferrets, salsa dancing and living in the moment.
Now that Greg Focker is in with his soon-to-be in-laws, Jack and Dina Byrnes, it looks like smooth sailing for him and his fiancée, Pam. But that's before Pam's parents meet Greg's parents, Bernie and Roz Focker. The hyper-relaxed Fockers and the tightly-wound Byrneses are woefully mismatched from the start, and no matter how hard Greg and Pam try, there is just no bringing their families together.