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Anna is a young widow who is finally getting on with her life after the death of her husband, Sean. Now engaged to be married, Anna meets a ten-year-old boy who tells her he is Sean reincarnated. Though his story is both unsettling and absurd, Anna can't get the boy out of her mind. And much to the concern of her fiancée, her increased contact with him leads her to question the choices she has made in her life.
Internationally famous oceanographer Steve Zissou (Bill Murray) and his crew—Team Zissou—set sail on an expedition to hunt down the mysterious, elusive, possibly non-existant Jaguar Shark that killed Zissou's partner during the documentary filming of their latest adventure. They are joined on their voyage by a young airline co-pilot who may or may not be Zissou's son (Owen Wilson), a beautiful journalist (Cate Blanchett) assigned to write a profile of Zissou, and Zissou's estranged wife and co-producer, Eleanor (Anjelica Huston). They face overwhelming complications including pirates, kidnapping, and bankruptcy.